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General citation for IQ-TREE 3:
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* T.K.F. Wong, N. Ly-Trong, H. Ren, H. Baños, A. Roger, E. Susko, C. Bielow, N. De Maio, N. Goldman, M.W. Hahn, G. Huttley, R. Lanfear, B.Q. Minh (2025)
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* Thomas K.F. Wong, Nhan Ly-Trong, Huaiyan Ren, Hector Banos, Andrew J. Roger, Edward Susko, Chris Bielow, Nicola De Maio, Nick Goldman, Matthew W. Hahn, Gavin Huttley, Robert Lanfear, Bui Quang Minh (2025)
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IQ-TREE 3: Phylogenomic Inference Software using Complex Evolutionary Models.
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*ecoevorxiv*. <https://ecoevorxiv.org/repository/view/8916/>
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*Submitted*. <https://ecoevorxiv.org/repository/view/8916/>
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Citation for IQ-TREE 2:
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When using MixtureFinder please cite:
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* B.Q. Minh, H.A. Schmidt, O. Chernomor, D. Schrempf, M.D. Woodhams, A. von Haeseler, R. Lanfear (2020)
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IQ-TREE 2: New models and efficient methods for phylogenetic inference in the genomic era.
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*Mol. Biol. Evol.*, 37:1530-1534. <https://doi.org/10.1093/molbev/msaa015>
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* Huaiyan Ren, Thomas KF Wong, Bui Quang Minh, Robert Lanfear (2025)
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MixtureFinder: Estimating DNA Mixture Models for Phylogenetic Analyses.
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*Mol. Biol. Evol.*, 42:msae264. <https://doi.org/10.1093/molbev/msae264>
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When using GTRpmix please cite:
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* Hector Banos, Thomas KF Wong, Justin Daneau, Edward Susko, Bui Quang Minh, Robert Lanfear, Matthew W Brown, Laura Eme, Andrew J Roger (2025)
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GTRpmix: A linked general-time reversible model for profile mixture models.
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*Mol. Biol. Evol.*, 92:msae174. <https://doi.org/10.1093/molbev/msae174>
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When using tree mixture models (MAST) please cite:
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When using CMAPLE please cite:
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* T.K.F. Wong, C. Cherryh, A.G. Rodrigo, M.W. Hahn, B.Q. Minh, R. Lanfear (2024)
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* Nhan Ly-Trong, Chris Bielow, Nicola De Maio, Bui Quang Minh (2024)
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CMAPLE: Efficient phylogenetic inference in the pandemic era.
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*Mol. Biol. Evol.*, 41:msae134. <https://doi.org/10.1093/molbev/msae134>
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When using the MAST model please cite:
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* Thomas KF Wong, Caitlin Cherryh, Allen G Rodrigo, Matthew W Hahn, Bui Quang Minh, Robert Lanfear (2024)
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MAST: Phylogenetic Inference with Mixtures Across Sites and Trees.
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_Syst. Biol._, in press. <https://doi.org/10.1093/sysbio/syae008>
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*Syst. Biol.*, 73:375–391. <https://doi.org/10.1093/sysbio/syae008>
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When computing concordance factors please cite:
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When using site/gene concordance factor please cite:
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* Y.K. Mo, R. Lanfear, M.W. Hahn, B.Q. Minh (2023)
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* Yu K Mo, Robert Lanfear, Matthew W Hahn, Bui Quang Minh (2023)
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Updated site concordance factors minimize effects of homoplasy and taxon sampling.
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_Bioinformatics_, 39:btac741. <https://doi.org/10.1093/bioinformatics/btac741>
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*Bioinformatics*, 39:btac741. <https://doi.org/10.1093/bioinformatics/btac741>
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When using AliSim to simulate alignments please cite:
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When using AliSim please cite:
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* N. Ly-Trong, G.M.J. Barca, B.Q. Minh (2023)
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* Nhan Ly-Trong, Giuseppe MJ Barca, Bui Quang Minh (2023)
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AliSim-HPC: parallel sequence simulator for phylogenetics.
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*Bioinformatics*, 39:btad540. <https://doi.org/10.1093/bioinformatics/btad540>
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When estimating amino-acid Q matrix please cite:
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When using QMaker for estimating protein Q matrix please cite:
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* Bui Quang Minh, Cuong Cao Dang, Le Sy Vinh, Robert Lanfear (2021)
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QMaker: Fast and Accurate Method to Estimate Empirical Models of Protein Evolution.
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*Syst. Biol.*, 70:1046–1060. <https://doi.org/10.1093/sysbio/syab010>
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When using nQMaker for estimating non-reversible Q matrix please cite:
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* B.Q. Minh, C. Cao Dang, L.S. Vinh, R. Lanfear (2021)
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QMaker: Fast and accurate method to estimate empirical models of protein evolution.
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_Syst. Biol._, 70:1046–1060. <https://doi.org/10.1093/sysbio/syab010>
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* Cuong Cao Dang, Bui Quang Minh, Hanon McShea, Joanna Masel, Jennifer E James, Le Sy Vinh, Robert Lanfear (2022)
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nQMaker: Estimating Time Nonreversible Amino Acid Substitution Models.
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*Syst. Biol.*, 71:1110–1123. <https://doi.org/10.1093/sysbio/syac007>
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When using the heterotachy GHOST model "+H" please cite:
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When using Rootstrap please cite:
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* S.M. Crotty, B.Q. Minh, N.G. Bean, B.R. Holland, J. Tuke, L.S. Jermiin, A. von Haeseler (2020)
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* Suha Naser-Khdour, Bui Quang Minh, Robert Lanfear (2022)
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Assessing Confidence in Root Placement on Phylogenies: An Empirical Study Using Nonreversible Models for Mammals.
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*Syst. Biol.*, 71:959–972. <https://doi.org/10.1093/sysbio/syab067>
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When using the GHOST \"+H\" model please cite:
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* Stephen M Crotty, Bui Quang Minh, Nigel G Bean, Barbara R Holland, Jonathan Tuke, Lars S Jermiin, Arndt Von Haeseler (2020)
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GHOST: Recovering Historical Signal from Heterotachously Evolved Sequence Alignments.
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_Syst. Biol._, 69:249-264. <https://doi.org/10.1093/sysbio/syz051>
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*Syst. Biol.*, 69:249-264. <https://doi.org/10.1093/sysbio/syz051>
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When using the tests of symmetry please cite:
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When using tests of symmetry please cite:
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* S. Naser-Khdour, B.Q. Minh, W. Zhang, E.A. Stone, R. Lanfear (2019)
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The Prevalence and Impact of Model Violations in Phylogenetic Analysis.
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* Suha Naser-Khdour, Bui Quang Minh, Wenqi Zhang, Eric A Stone, Robert Lanfear (2019)
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The Prevalence and Impact of Model Violations in Phylogenetic Analysis.
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*Genome Biol. Evol.*, 11:3341-3352. <https://doi.org/10.1093/gbe/evz193>
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When using polymorphism-aware models please cite:
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* D. Schrempf, B.Q. Minh, A. von Haeseler, C. Kosiol (2019)
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Polymorphism-aware species trees with advanced mutation models, bootstrap, and rate heterogeneity.
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*Mol. Biol. Evol.*, 36:1294–1301. <https://doi.org/10.1093/molbev/msz043>
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When using ModelFinder please cite:
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For the ultrafast bootstrap (UFBoot) please cite:
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* Subha Kalyaanamoorthy, Bui Quang Minh, Thomas KF Wong, Arndt von Haeseler, and Lars S Jermiin (2017)
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ModelFinder: Fast model selection for accurate phylogenetic estimates.
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*Nat. Methods*, 14:587–589. <https://doi.org/10.1038/nmeth.4285>
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* D.T. Hoang, O. Chernomor, A. von Haeseler, B.Q. Minh, and L.S. Vinh (2018)
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UFBoot2: Improving the ultrafast bootstrap approximation.
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*Mol. Biol. Evol.*, 35:518–522. <https://doi.org/10.1093/molbev/msx281>
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When using polymorphism-aware models please cite:
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* Dominik Schrempf, Bui Quang Minh, Arndt von Haeseler, Carolin Kosiol (2019)
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Polymorphism-Aware Species Trees with Advanced Mutation Models, Bootstrap, and Rate Heterogeneity.
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*Mol. Biol. Evol.*, 36:1294–1301. <https://doi.org/10.1093/molbev/msz043>
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When using posterior mean site frequency model (PMSF) please cite:
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When using site-specific frequency model please cite:
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* H.C. Wang, B.Q. Minh, S. Susko, A.J. Roger (2018)
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Modeling site heterogeneity with posterior mean site frequency profiles
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accelerates accurate phylogenomic estimation.
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* Huai-Chun Wang, Edward Susko, Bui Quang Minh, and Andrew J. Roger (2018)
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Modeling site heterogeneity with posterior mean site frequency profiles accelerates accurate phylogenomic estimation.
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*Syst. Biol.*, 67:216–235. <https://doi.org/10.1093/sysbio/syx068>
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When using ModelFinder please cite:
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* S. Kalyaanamoorthy, B.Q. Minh, T.K.F. Wong, A. von Haeseler, L.S. Jermiin (2017)
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ModelFinder: Fast model selection for accurate phylogenetic estimates.
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*Nat. Methods*, 14:587-589. <https://doi.org/10.1038/nmeth.4285>
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When using ultrafast bootstrap (UFBoot) please cite:
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* Diep Thi Hoang, Olga Chernomor, Arndt von Haeseler, Bui Quang Minh, and Le Sy Vinh (2018)
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UFBoot2: Improving the ultrafast bootstrap approximation.
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*Mol. Biol. Evol.*, 35:518–522. <https://doi.org/10.1093/molbev/msx281>
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When using partition models please cite:
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* O. Chernomor, A. von Haeseler, B.Q. Minh (2016)
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* Olga Chernomor, Arndt von Haeseler, and Bui Quang Minh (2016)
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Terrace aware data structure for phylogenomic inference from supermatrices.
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*Syst. Biol.*, 65:997-1008. <https://doi.org/10.1093/sysbio/syw037>
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When using terrace analysis please cite:
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* Biczok R, Bozsoky P, Eisenmann P, Ernst J, Ribizel T, Scholz F, Trefzer A, Weber F, Hamann M, Stamatakis A. (2018)
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Two C++ libraries for counting trees on a phylogenetic terrace.
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*Bioinformatics*, 34:3399–3401. <https://doi.org/10.1093/bioinformatics/bty384>
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When using least square dating (LSD) please cite:
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* Thu-Hien To, Matthieu Jung, Samantha Lycett, Olivier Gascuel (2016)
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Fast dating using least-squares criteria and algorithms.
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*Syst. Biol.*, 65:82-97. <https://doi.org/10.1093/sysbio/syv068>
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When using IQ-TREE web server please cite:
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* J. Trifinopoulos, L.-T. Nguyen, A. von Haeseler, B.Q. Minh (2016)
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W-IQ-TREE: a fast online phylogenetic tool for maximum likelihood analysis.
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*Nucleic Acids Res.*, 44:W232-W235. <https://doi.org/10.1093/nar/gkw256>
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When using IQ-TREE 2 please cite:
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* B.Q. Minh, H.A. Schmidt, O. Chernomor, D. Schrempf, M.D. Woodhams, A. von Haeseler, R. Lanfear (2020)
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IQ-TREE 2: New models and efficient methods for phylogenetic inference in the genomic era.
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*Mol. Biol. Evol.*, 37:1530-1534. <https://doi.org/10.1093/molbev/msaa015>
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When using IQ-TREE version 1 please cite:
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* L. Nguyen, H.A. Schmidt, A. von Haeseler, B.Q. Minh (2015)
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IQ-TREE: A Fast and Effective Stochastic Algorithm for Estimating Maximum-Likelihood Phylogenies.
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_Mol. Biol. and Evol._, 32:268-274. <https://doi.org/10.1093/molbev/msu300>
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*Mol. Biol. Evol.*, 32:268-274. <https://doi.org/10.1093/molbev/msu300>
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Development team

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